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sc-PDB

An Annotated Database of Druggable Binding Sites from the Protein DataBank

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Distribution of Interaction pattern similarity measured by Grim
You can zoom onto the graph by using the mouse to make a selection

Binding Modes are compared using Grim.
For more information, please see the following publication: Desaphy J. et al. Encoding Protein-Ligand Interaction Patterns in Fingerprints and Graphs J. Chem. Inf. Model., 2013, 53 (3), pp 623-637
Binding modes are considered as similar when the similarity value is greater than 0.65


Reference Protein Data Bank Entry :
PDB ID HET Uniprot Name EC Number
4kln AGS Transitional endoplasmic reticulum ATPase 3.6.4.6

Complex with similar interaction patterns:

PDB ID HET Uniprot Name EC Number Binding Mode
Similarity		 Align
4kln AGSTransitional endoplasmic reticulum ATPase 3.6.4.6 1.062
4ko8 AGSTransitional endoplasmic reticulum ATPase 3.6.4.6 0.891
3hu1 AGSTransitional endoplasmic reticulum ATPase 3.6.4.6 0.884
3hu3 AGSTransitional endoplasmic reticulum ATPase 3.6.4.6 0.869
3hu2 AGSTransitional endoplasmic reticulum ATPase 3.6.4.6 0.837
3d8b ADPFidgetin-like protein 1 3.6.4 0.793
1gil GSPGuanine nucleotide-binding protein G(i) subunit alpha-1 / 0.785
1gwn GTPRho-related GTP-binding protein RhoE / 0.772
3tkl GTPRas-related protein Rab-1A / 0.766
2cls GTPRho-related GTP-binding protein Rho6 / 0.764
1cxz GSPTransforming protein RhoA / 0.761
1ytm ATPPhosphoenolpyruvate carboxykinase (ATP) / 0.760
3vx4 ATPPutative ABC transporter, ATP-binding protein ComA / 0.758
2olq ATPPhosphoenolpyruvate carboxykinase (ATP) / 0.755
5dha GNPGTP-binding nuclear protein Ran / 0.754
4kvg GTPRas-related protein Rap-1A / 0.752
1cul GSPGuanine nucleotide-binding protein G(s) subunit alpha isoforms short / 0.751
2cl6 CAGGTPase HRas / 0.751
4tl7 ATPCircadian clock protein kinase KaiC 2.7.11.1 0.749
1e96 GTPRas-related C3 botulinum toxin substrate 1 / 0.746
4dvg GSPRho family GTPase / 0.746
1wa5 GTPGTP-binding nuclear protein Ran / 0.743
1mh1 GNPRas-related C3 botulinum toxin substrate 1 / 0.742
1ayl ATPPhosphoenolpyruvate carboxykinase (ATP) / 0.741
4dso GSPGTPase KRas / 0.741
4xsg GSPTransforming protein RhoA / 0.741
2gvd GSPGuanine nucleotide-binding protein G(s) subunit alpha isoforms short / 0.740
3bos CDPRegulatory inactivation of DnaA Hda protein / 0.740
3vhx GTPADP-ribosylation factor 6 / 0.740
4jsy GTPMetallophosphoesterase / 0.740
1s3s ADPTransitional endoplasmic reticulum ATPase 3.6.4.6 0.738
1cs4 GSPGuanine nucleotide-binding protein G(s) subunit alpha isoforms short / 0.737
4tl8 AGSCircadian clock protein kinase KaiC 2.7.11.1 0.736
3tgp GNPGTPase HRas / 0.735
4luc GDPGTPase KRas / 0.734
4tld AGSCircadian clock protein kinase KaiC 2.7.11.1 0.734
1os1 ATPPhosphoenolpyruvate carboxykinase (ATP) / 0.733
2c5l GTPGTPase HRas / 0.731
4idn GNPAtlastin-1 3.6.5 0.731
1fmw ATPMyosin-2 heavy chain / 0.729
1jx2 ADPMyosin-2 heavy chain / 0.729
1l8q ADPChromosomal replication initiator protein DnaA / 0.727
4lrw GDPGTPase KRas / 0.727
2gcp GSPRho-related GTP-binding protein RhoC / 0.726
1bmf ADPATP synthase subunit alpha, mitochondrial / 0.725
1bmf ADPATP synthase subunit beta, mitochondrial 3.6.3.14 0.725
1gia GSPGuanine nucleotide-binding protein G(i) subunit alpha-1 / 0.724
4gzl GNPRas-related C3 botulinum toxin substrate 1 / 0.724
1xmv ADPProtein RecA / 0.723
4tle AGSCircadian clock protein kinase KaiC 2.7.11.1 0.723
1t91 GTPRas-related protein Rab-7a / 0.722
1wdt GTPElongation factor G (EF-G-2) / 0.722
2xtz GSPGuanine nucleotide-binding protein alpha-1 subunit / 0.722
3cx8 GSPGuanine nucleotide-binding protein subunit alpha-13 / 0.722
2py7 ATPPhosphoenolpyruvate carboxykinase (ATP) / 0.721
5c46 GSPRas-related protein Rab-11A / 0.721
2j5x GSPADP-ribosylation factor 6 / 0.720
4xoi GTPTransforming protein RhoA / 0.720
3c16 GSPGuanine nucleotide-binding protein G(s) subunit alpha isoforms short / 0.719
3cob ADPKinesin heavy chain-like protein / 0.718
3kz1 GSPTransforming protein RhoA / 0.718
3zfd ANPChromosome-associated kinesin KIF4 / 0.718
4tlc AGSCircadian clock protein kinase KaiC 2.7.11.1 0.718
1f5n GNPGuanylate-binding protein 1 / 0.717
3myk ANPMyosin-2 heavy chain / 0.715
5bn3 ADPV-type ATP synthase alpha chain / 0.714
5bn3 ADPNEQ263 / 0.714
1mmg AGSMyosin-2 heavy chain / 0.713
1yu9 GNPRas-related protein Rab-4A / 0.713
3qbt GNPRas-related protein Rab-8A / 0.713
3tjz GNPADP-ribosylation factor 1 / 0.713
4dxa GSPRas-related protein Rap-1b / 0.713
4qxa GTPRas-related protein Rab-9A / 0.713
4ww0 ADPATP-dependent zinc metalloprotease FtsH / 0.713
5irn ADPNucleotide binding oligomerization domain containing 2 / 0.713
4a1z ADPKinesin-like protein KIF11 / 0.712
4qm6 GTPMetallophosphoesterase / 0.712
4mit GTPRho-related protein racC / 0.711
1w1w AGSStructural maintenance of chromosomes protein 1 / 0.710
1aq2 ATPPhosphoenolpyruvate carboxykinase (ATP) / 0.709
2qen ADPWalker-Type ATPase / 0.709
4kod ADPTransitional endoplasmic reticulum ATPase 3.6.4.6 0.709
4v02 ATPSite-determining protein / 0.709
1azt GSPGuanine nucleotide-binding protein G(s) subunit alpha isoforms short / 0.708
1e1q ADPATP synthase subunit beta, mitochondrial 3.6.3.14 0.707
1svm ATPLarge T antigen 3.6.4 0.707
4o25 GTPGTP-binding protein Rheb / 0.707
1w5t ANPORC1-type DNA replication protein 2 / 0.705
3k1j ADPATP-dependent protease Lon / 0.705
4d0l GSPRas-related protein Rab-11A / 0.705
1k5d GNPGTP-binding nuclear protein Ran / 0.704
1khe GCPPhosphoenolpyruvate carboxykinase, cytosolic [GTP] 4.1.1.32 0.704
1e79 ATPATP synthase subunit alpha, mitochondrial / 0.703
1e79 ATPATP synthase subunit beta, mitochondrial 3.6.3.14 0.703
1ny5 ADPTranscriptional regulator (NtrC family) / 0.703
3b1x GNPFerrous iron transport protein B / 0.703
3m0e ATPTranscriptional regulator (NtrC family) / 0.703
4cvl ACPUDP-N-acetylmuramoyl-tripeptide--D-alanyl-D-alanine ligase / 0.703
1a2b GSPTransforming protein RhoA / 0.702
2x7e ADPKinesin-like protein KIF11 / 0.702
2xae ADPKinesin-like protein KIF11 / 0.702
3fwy ADPLight-independent protochlorophyllide reductase iron-sulfur ATP-binding protein / 0.702
1khb GCPPhosphoenolpyruvate carboxykinase, cytosolic [GTP] 4.1.1.32 0.701
3ftq GNPSeptin-2 / 0.701
4hdo GNPRas-related protein Rap-1b / 0.701
4hdq GNPRas-related protein Rap-1b / 0.701
2x2r ADPKinesin-like protein KIF11 / 0.700
2ce2 GDPGTPase HRas / 0.699
2g1q ADPKinesin-like protein KIF11 / 0.699
2jj2 ADPATP synthase subunit alpha, mitochondrial / 0.699
2jj2 ADPATP synthase subunit beta, mitochondrial 3.6.3.14 0.699
4hse ADPChaperone protein ClpB / 0.699
2bme GNPRas-related protein Rab-4A / 0.698
3i5x ANPATP-dependent RNA helicase MSS116, mitochondrial 3.6.4.13 0.698
1in5 ADPHolliday junction ATP-dependent DNA helicase RuvB / 0.697
2pg2 ADPKinesin-like protein KIF11 / 0.697
4epw GDPGTPase KRas / 0.697
4pjk ADPMyosin-2 heavy chain / 0.697
2q3f GNPRas-related GTP-binding protein D / 0.696
4a51 ADPKinesin-like protein KIF11 / 0.696
4d81 ADPAAA ATPase, central domain protein / 0.696
4lpk GDPGTPase KRas / 0.696
1e79 ADPATP synthase subunit alpha, mitochondrial / 0.695
1e79 ADPATP synthase subunit beta, mitochondrial 3.6.3.14 0.695
2o8b ADPDNA mismatch repair protein Msh2 / 0.695
4pfp ADPUnconventional myosin-VI / 0.695
1fnn ADPORC1-type DNA replication protein / 0.694
1mma ADPMyosin-2 heavy chain / 0.694
2bif ANP6-phosphofructo-2-kinase/fructose-2,6-bisphosphatase 4 2.7.1.105 0.694
1sdm ADPKinesin heavy chain-like protein / 0.693
1u0h GSPGuanine nucleotide-binding protein G(s) subunit alpha isoforms short / 0.693
2r9v ATPATP synthase subunit alpha / 0.693
4db1 ANPMyosin-7 / 0.693
4pjj ADPUnconventional myosin-VI / 0.693
2dpx GDPGTP-binding protein RAD / 0.692
5dac AGSUncharacterized protein / 0.692
3q5d GDPAtlastin-1 3.6.5 0.691
4zca ADPKinesin-like protein KIF11 / 0.691
2rgg GNPGTPase HRas / 0.690
2y65 ADPKinesin heavy chain / 0.690
2zfi ADPKinesin-like protein KIF1A / 0.690
2fju GSPRas-related C3 botulinum toxin substrate 1 / 0.689
3x1w GDPRas-related protein Rap-1b / 0.689
5aun ADPIron-sulfur cluster carrier protein / 0.689
1yrs ADPKinesin-like protein KIF11 / 0.688
3lbi GNPGTPase HRas / 0.688
3oee ANPATP synthase subunit alpha, mitochondrial / 0.688
3oee ANPATP synthase subunit beta, mitochondrial 3.6.3.14 0.688
4cvm ANPUDP-N-acetylmuramoyl-tripeptide--D-alanyl-D-alanine ligase / 0.688
3x1z GNPRas-related protein Rap-1b / 0.687
2uyi ADPKinesin-like protein KIF11 / 0.686
3rab GNPRas-related protein Rab-3A / 0.686
3w3z GTPGTP-binding nuclear protein Ran / 0.686
4oi2 ADPProtein clpf-1 / 0.686
1w5t ADPORC1-type DNA replication protein 2 / 0.685
2jlr ANPGenome polyprotein 3.4.21.91 0.685
3oeh ANPATP synthase subunit alpha, mitochondrial / 0.685
4obe GDPGTPase KRas / 0.685
4ohv ANPProtein clpf-1 / 0.684
4tlb AGSCircadian clock protein kinase KaiC 2.7.11.1 0.684
1z0f GDPRas-related protein Rab-14 / 0.683
3ddc GNPGTPase HRas / 0.683
4ql3 GDPGTPase KRas / 0.683
1oix GDPRas-related protein Rab-11A / 0.682
3tso GNPRas-related protein Rab-25 / 0.682
4c30 ANPDNA helicase / 0.682
4ldj GDPGTPase KRas / 0.682
4pao GSPGuanine nucleotide-binding protein G(i) subunit alpha-1 / 0.682
1d5c GDPGTPase (Rab6) / 0.681
1kmq GNPTransforming protein RhoA / 0.681
1x1r GDPRas-related protein M-Ras / 0.681
2hld ANPATP synthase subunit alpha, mitochondrial / 0.681
2hld ANPATP synthase subunit beta, mitochondrial 3.6.3.14 0.681
4dsu GDPGTPase KRas / 0.681
4q02 GDPGTPase KRas / 0.681
4xvu ATPATPase GET3 / 0.681
2olr ATPPhosphoenolpyruvate carboxykinase (ATP) / 0.680
3cph GDPRas-related protein SEC4 / 0.680
3kuc GDPRas-related protein Rap-1A / 0.680
3rs4 GNPGTPase HRas / 0.680
3rs7 GNPGTPase HRas / 0.680
3rsl GNPGTPase HRas / 0.680
4a28 ADPKinesin-like protein KIF11 / 0.680
4bzq ADPBifunctional enzyme CysN/CysC 2.7.1.25 0.680
5cyo GDPSeptin-9 / 0.680
5ftc ADPTPR domain protein / 0.680
1fp6 ADPNitrogenase iron protein 1 1.18.6.1 0.679
1m7b GTPRho-related GTP-binding protein RhoE / 0.679
3ea5 GDPGTP-binding nuclear protein Ran / 0.679
3oiu GNPGTPase HRas / 0.678
3pcr GTPADP-ribosylation factor 6 / 0.678
4dlv GNPGTPase HRas / 0.678
4g0n GNPGTPase HRas / 0.678
4goj GNPADP-ribosylation factor-like protein 3 / 0.678
4l9w GNPGTPase HRas / 0.678
4nh0 ATPESX secretion system protein EccC / 0.678
4yqf GDPSeptin-9 / 0.678
5i67 GDPPhosphoenolpyruvate carboxykinase [GTP] / 0.678
1h65 GDPTranslocase of chloroplast 34 3.6.5 0.677
1h8h ATPATP synthase subunit alpha, mitochondrial / 0.677
1h8h ATPATP synthase subunit beta, mitochondrial 3.6.3.14 0.677
3l8y GNPGTPase HRas / 0.677
3oiw GNPGTPase HRas / 0.677
3v4f GNPGTPase HRas / 0.677
4dls GNPGTPase HRas / 0.677
4dlt GNPGTPase HRas / 0.677
1v8k ANPKinesin-like protein KIF2C / 0.676
1wbb ADPDNA mismatch repair protein MutS / 0.676
4dlz GNPGTPase HRas / 0.676
4tl6 ANPCircadian clock protein kinase KaiC 2.7.11.1 0.676
4z54 GDPNeuronal-specific septin-3 / 0.676
1z0d GDPRas-related protein Rab-5C / 0.675
1e1q ANPATP synthase subunit alpha, mitochondrial / 0.674
1e1q ANPATP synthase subunit beta, mitochondrial 3.6.3.14 0.674
2rgb GNPGTPase HRas / 0.674
3a4l ANPL-seryl-tRNA(Sec) kinase 2.7.1.164 0.674
3adc ANPL-seryl-tRNA(Sec) kinase 2.7.1.164 0.674
4pk4 ADPUnconventional myosin-VI / 0.674
4ymu ATPABC-type polar amino acid transport system, ATPase component / 0.674
1phk ATPPhosphorylase b kinase gamma catalytic chain, skeletal muscle/heart isoform 2.7.11.19 0.673
1r0z ATPCystic fibrosis transmembrane conductance regulator / 0.673
1s1c GNPTransforming protein RhoA / 0.673
2fg5 GNPRas-related protein Rab-31 / 0.673
2rgx AP5Adenylate kinase / 0.673
3rs5 GNPGTPase HRas / 0.673
4cf7 ADPAdenylate kinase / 0.673
4xvq GNPGTPase HRas / 0.673
1n6l GTPRas-related protein Rab-5A / 0.672
1sxj AGSReplication factor C subunit 1 / 0.672
1uad GNPRas-related protein Ral-A / 0.672
2dr3 ADPUPF0273 protein PH0284 / 0.672
2jj1 ADPATP synthase subunit alpha, mitochondrial / 0.672
2jj1 ADPATP synthase subunit beta, mitochondrial 3.6.3.14 0.672
3rs0 GNPGTPase HRas / 0.672
3zia ATPATP synthase subunit alpha, mitochondrial / 0.672
5b2z GNPGTPase HRas / 0.672
1am4 GNPCell division control protein 42 homolog / 0.671
1nsf ATPVesicle-fusing ATPase 3.6.4.6 0.671
2aky AP5Adenylate kinase / 0.671
2jj1 ANPATP synthase subunit alpha, mitochondrial / 0.671
2rgc GNPGTPase HRas / 0.671
4cid ANPEH domain-containing protein 2 / 0.671
4dlx GNPGTPase HRas / 0.671
2ck3 ANPATP synthase subunit alpha, mitochondrial / 0.670
2ck3 ANPATP synthase subunit beta, mitochondrial 3.6.3.14 0.670
2ffq GSPRas-related protein Rab-6B / 0.670
3e5h GNPRas-related protein Rab-28 / 0.670
3kko GNPRas-related protein M-Ras / 0.670
3l9h ADPKinesin-like protein KIF11 / 0.670
4ap0 ADPKinesin-like protein KIF11 / 0.670
4asu ADPATP synthase subunit alpha, mitochondrial / 0.670
4dlw GNPGTPase HRas / 0.670
4epx GDPGTPase KRas / 0.670
4xvr GNPGTPase HRas / 0.670
2rge GNPGTPase HRas / 0.669
3dzd ADPTranscriptional regulator (NtrC family) / 0.669
3k8y GNPGTPase HRas / 0.669
3kud GDPGTPase HRas / 0.669
3law GNPRas-related protein Rab-7a / 0.669
3mjh GTPRas-related protein Rab-5A / 0.669
3wyf GTPGTP-binding nuclear protein / 0.669
4g5q GDPGuanine nucleotide-binding protein G(i) subunit alpha-1 / 0.669
1got GDPGuanine nucleotide-binding protein G(t) subunit alpha-1 / 0.668
3k9l GNPGTPase HRas / 0.668
4lhv GDPRas-related protein Rab-8A / 0.668
1bmf ANPATP synthase subunit alpha, mitochondrial / 0.667
1bmf ANPATP synthase subunit beta, mitochondrial 3.6.3.14 0.667
1p3j AP5Adenylate kinase / 0.667
1svl ADPLarge T antigen 3.6.4 0.667
1tu3 GNPRas-related protein Rab-5A / 0.667
2fl2 ADPKinesin-like protein KIF11 / 0.667
3bc1 GNPRas-related protein Rab-27A / 0.667
4lv6 GDPGTPase KRas / 0.667
1ank ANPAdenylate kinase / 0.666
2fna ADPUncharacterized protein / 0.666
3dkv AP5Adenylate kinase / 0.666
3oes GNPGTPase RhebL1 / 0.666
4uj3 GNPRas-related protein Rab-11A / 0.666
5jlj GNPGTP-binding nuclear protein Ran / 0.666
1xef ATPAlpha-hemolysin translocation ATP-binding protein HlyB / 0.665
1zd9 GDPADP-ribosylation factor-like protein 8A / 0.665
2xsz ATPRuvB-like 2 3.6.4.12 0.665
3c41 ANPAmino acid ABC transporter / 0.665
3th5 GNPRas-related C3 botulinum toxin substrate 1 / 0.665
4js0 GNPCell division control protein 42 homolog / 0.665
1um8 ADPATP-dependent Clp protease ATP-binding subunit ClpX / 0.664
1zc4 GNPRas-related protein Ral-A / 0.664
2uz3 TTPThymidine kinase / 0.664
2v7q ADPATP synthase subunit beta, mitochondrial 3.6.3.14 0.664
3fvq ATPFe(3+) ions import ATP-binding protein FbpC / 0.664
3sbd GNPRas-related C3 botulinum toxin substrate 1 / 0.664
4tyq AP5Adenylate kinase / 0.664
1svw GTPProbable GTP-binding protein EngB / 0.663
2qme GCPRas-related C3 botulinum toxin substrate 3 / 0.663
4lhw GNPRas-related protein Rab-8A / 0.663
4tq9 GDPGTPase KRas / 0.663
1w0k ADPATP synthase subunit alpha, mitochondrial / 0.662
1w0k ADPATP synthase subunit beta, mitochondrial 3.6.3.14 0.662
3eg5 GNPCell division control protein 42 homolog / 0.662
3lbn GNPGTPase HRas / 0.662
4itr GDPCell division control protein 42 homolog / 0.662
4ohx ADPProtein clpf-1 / 0.662
4q03 GDPGTPase KRas / 0.662
4wa7 GDPGTPase KRas / 0.662
1b7t ADPMyosin heavy chain, striated muscle / 0.661
1njf ADPDNA polymerase III subunit tau 2.7.7.7 0.661
1ohh ANPATP synthase subunit alpha, mitochondrial / 0.661
1ohh ANPATP synthase subunit beta, mitochondrial 3.6.3.14 0.661
2qtv GNPSmall COPII coat GTPase SAR1 3.6.5 0.661
3mle ADPATP-dependent dethiobiotin synthetase BioD 6.3.3.3 0.661
4gp6 ADPMetallophosphoesterase / 0.661
4l16 ADPFidgetin-like protein 1 3.6.4 0.661
1i4l GDPRas-related C3 botulinum toxin substrate 1 / 0.660
1h8h ADPATP synthase subunit alpha, mitochondrial / 0.659
1h8h ADPATP synthase subunit beta, mitochondrial 3.6.3.14 0.659
3dl0 AP5Adenylate kinase / 0.659
5dif GNPGTP-binding nuclear protein Ran / 0.659
1w7i ADPUnconventional myosin-Va / 0.658
2ov2 GCPRas-related C3 botulinum toxin substrate 3 / 0.658
4ys0 ADPProtein translocase subunit SecA / 0.658
1ake AP5Adenylate kinase / 0.657
1yzq GNPRas-related protein Rab-6A / 0.657
1z2a GDPRas-related protein Rab-23 / 0.657
2j7p GNPSignal recognition particle protein / 0.657
2j7p GNPSignal recognition particle receptor FtsY / 0.657
2oap ANPType II secretion system protein (GspE-2) / 0.657
3hqd ANPKinesin-like protein KIF11 / 0.657
4nm7 ADPGlycogen synthase kinase-3 beta 2.7.11.26 0.657
1gp2 GDPGuanine nucleotide-binding protein G(i) subunit alpha-1 / 0.656
2bc9 GNPGuanylate-binding protein 1 / 0.656
2iut AGSDNA translocase FtsK / 0.656
4c0b ATPmRNA cleavage and polyadenylation factor CLP1 / 0.656
2a5y ATPCell death protein 4 / 0.655
2j0s ANPEukaryotic initiation factor 4A-III 3.6.4.13 0.655
3pit GNPRas-related protein M-Ras / 0.655
4g01 GDPRas-related protein RABF2b / 0.655
4tl9 AGSCircadian clock protein kinase KaiC 2.7.11.1 0.655
4xj4 3ATCyclic GMP-AMP synthase / 0.655
1ow3 GDPTransforming protein RhoA / 0.654
2cea ADPATP-dependent zinc metalloprotease FtsH / 0.654
2hf8 GSPProbable hydrogenase nickel incorporation protein HypB / 0.654
2x3j ATPAcsD / 0.654
3i3s GNPGTPase HRas / 0.654
4jlo ADPAdenylate kinase / 0.654
2v7q ATPATP synthase subunit alpha, mitochondrial / 0.653
2v7q ATPATP synthase subunit beta, mitochondrial 3.6.3.14 0.653
4jld ADPAdenylate kinase / 0.653
4rkf GNPRas-related protein Rab-3 / 0.653
1gqy ACPUDP-N-acetylmuramate--L-alanine ligase 6.3.2.8 0.652
2qaj AP5Adenylate kinase / 0.652
2w2x GSPRas-related C3 botulinum toxin substrate 2 / 0.652
3zvm ADPBifunctional polynucleotide phosphatase/kinase 2.7.1.78 0.652
5di3 GNPADP-ribosylation factor-like protein 3 / 0.652
2dwp ACP6-phosphofructo-2-kinase/fructose-2,6-bisphosphatase 3 2.7.1.105 0.651
3c14 GSPGuanine nucleotide-binding protein G(s) subunit alpha isoforms short / 0.650